The L1 protein which coded by the late gene L1 of HPV18 is the main antigen which induces the neutralization antibody.

其晚期基因L1结构蛋白是刺激机体产生中和抗体的主要抗原。

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Faced with this dead end twenty years ago, the late Gene Upshaw chose a different path.

FORBES: How The Decertification Option Fits Into Players' Plan to Fight NFL

They feel the union is angling for the strategy employed by the NFLPA in the 1992-93 under the late Gene Upshaw , a strategy that led to many gains for NFL player including, for the first time, free agency.

FORBES: NFL Labor Pains: An Issue Of Trust

Dave Pear and Gene Upshaw were teammates in the late 1970s with the Oakland Raiders, you know, and Gene Upshaw is very upfront about saying, you know, he knew Dave Pear was injured there.

NPR: Former NFL Players Unable to Pay Medical Bills